01 · NCERT §12 (introduction)
Ecosystem: meaning and types
- An ecosystem is a functional unit of nature in which living organisms interact with each other and with their physical surroundings.
- An ecosystem can be as small as a pond or as large as a whole forest or sea.
- Many ecologists treat the whole biosphere as one global ecosystem, made up of all the local ecosystems on Earth.
- Since the biosphere is too large and complex to study at once, it is divided into two basic kinds: terrestrial and aquatic.
- Terrestrial examples: forest, grassland, desert. Aquatic examples: pond, lake, wetland, river, estuary.
- Crop fields and an aquarium count as man-made ecosystems.
- The chapter follows the input (productivity), the transfer of energy (food chains and webs) and the output (degradation and loss of energy).
02 · NCERT §12.1
Structure and function
- An ecosystem has abiotic (non-living) and biotic (living) components. Their interaction gives each type of ecosystem a characteristic physical structure.
- Identifying and counting the plants and animals present tells us the species composition of an ecosystem.
- Stratification is the vertical distribution of species at different levels.
- In a forest, trees form the top layer, shrubs the second, and herbs and grasses the bottom layers.
- The two chief features of an ecosystem's structure are its species composition and its stratification.
- An ecosystem works as a unit through four functional aspects: productivity, decomposition, energy flow and nutrient cycling.
03 · NCERT §12.1
The pond ecosystem
- A pond is a shallow water body and a fairly self-sustaining, simple example that shows all four basic functions of an ecosystem.
- Abiotic component: the water, the inorganic and organic matter dissolved in it, and the nutrient-rich soil on the pond floor.
- Solar input, the temperature cycle, day length and other climatic conditions regulate how fast the whole pond functions.
- Autotrophs (producers): phytoplankton, some algae, and plants that float, stay submerged or grow along the margins.
- Consumers: zooplankton, and free-swimming and bottom-dwelling forms.
- Decomposers: fungi, bacteria and flagellates, most abundant at the bottom of the pond.
- The pond does what every ecosystem does: autotrophs turn inorganic into organic matter using solar energy, heterotrophs consume the autotrophs, and dead matter is decomposed and mineralised for the autotrophs to reuse.
- Energy moves in one direction towards higher trophic levels and is lost as heat to the surroundings.
04 · NCERT §12.2
Productivity
- Every ecosystem needs a steady supply of solar energy to keep working.
- Primary production is the organic matter (biomass) that plants make by photosynthesis on a unit of area over a stated period. It is expressed as weight (g m⁻²) or energy (kcal m⁻²).
- Productivity is the rate of biomass production, expressed as g m⁻² yr⁻¹ or kcal m⁻² yr⁻¹, so that different ecosystems can be compared.
- Gross primary productivity (GPP) is the rate at which organic matter is made in photosynthesis. Plants use a considerable part of it in respiration.
- Net primary productivity: GPP − R = NPP, where R is the respiration loss. NPP is the biomass available to heterotrophs (herbivores and decomposers).
- Secondary productivity is the rate at which consumers form new organic matter.
- Primary productivity depends on the plant species of an area, environmental factors, nutrient availability and the photosynthetic capacity of plants, so it differs between ecosystems.
- The annual NPP of the whole biosphere is about 170 billion tons (dry weight) of organic matter.
- Oceans cover about 70 per cent of the surface but produce only 55 billion tons of it; the rest is produced on land.
05 · NCERT §12.3
Decomposition
- Decomposers break complex organic matter down into inorganic substances such as carbon dioxide, water and nutrients. This is decomposition.
- The earthworm is called the farmer's friend because it helps break down organic matter and loosens the soil.
- Detritus, the raw material, is made of dead plant remains (leaves, bark, flowers) and dead animal remains, including fecal matter.
- Fragmentation: detritivores such as the earthworm break detritus into smaller particles.
- Leaching: water-soluble inorganic nutrients move down into the soil horizon and are precipitated as unavailable salts.
- Catabolism: enzymes of bacteria and fungi break detritus down to simpler inorganic compounds. These three steps run at the same time on the detritus.
- Humification forms humus, a dark, amorphous substance that resists microbes and decomposes very slowly. Being colloidal, it acts as a nutrient reservoir.
- Mineralisation: some microbes degrade the humus further and release inorganic nutrients.
- Decomposition mostly needs oxygen. It is slower when detritus is rich in lignin and chitin, and faster when it is rich in nitrogen and water-soluble substances such as sugars.
- Of the climatic factors, temperature and soil moisture matter most, acting through soil microbes. Warm, moist conditions speed it up; low temperature and anaerobiosis slow it, so organic matter builds up.
06 · NCERT §12.4
Energy flow
- Apart from deep-sea hydrothermal ecosystems, every ecosystem on Earth runs on energy from the sun.
- Less than 50 per cent of the incident solar radiation is photosynthetically active radiation (PAR).
- Of this PAR, plants trap just 2-10 per cent, yet that small share keeps the whole living world going.
- All organisms depend on producers for food, directly or indirectly, so energy flows one way: from the sun to producers and then to consumers.
- Ecosystems obey the second law of thermodynamics. They need a constant energy supply to build the molecules they need and to counter the universal trend towards disorder.
- Producers in terrestrial ecosystems are mainly herbaceous and woody plants; in aquatic ecosystems they are phytoplankton, algae and higher plants.
- Energy trapped in an organism does not stay there for ever. It passes to a consumer, or the organism dies, and death starts the detritus food chain.
07 · NCERT §12.4
Food chains and food webs
- Consumers (heterotrophs) depend on plants directly or indirectly for food.
- Primary consumers eat producers, so they are herbivores. Examples: insects, birds and mammals on land; molluscs in water.
- Secondary consumers eat herbivores; they are primary carnivores. Animals that eat primary carnivores are secondary carnivores (tertiary consumers).
- A simple grazing food chain (GFC): grass (producer) → goat (primary consumer) → man (secondary consumer).
- The detritus food chain (DFC) starts from dead organic material. Its decomposers, mainly fungi and bacteria, are heterotrophs also called saprotrophs.
- Decomposers secrete digestive enzymes that break dead and waste materials into simple inorganic materials, which they then absorb.
- In aquatic ecosystems the GFC is the main channel of energy flow. On land, a much larger share of energy flows through the DFC than through the GFC.
- The DFC links with the GFC at some levels: some DFC organisms are eaten by GFC animals, and some animals, such as cockroaches and crows, are omnivores. These links make a food web.
08 · NCERT §12.4
Trophic levels and the 10 per cent law
- An organism's trophic level is its place in the food chain, based on its source of food.
- Producers form the first trophic level, herbivores (primary consumers) the second and carnivores (secondary consumers) the third.
- The amount of energy decreases at each successive trophic level.
- When any organism dies it becomes detritus, dead biomass that is an energy source for decomposers.
- The standing crop is the mass of living material at a trophic level at a given time, measured as biomass or as number per unit area.
- Biomass is given as fresh or dry weight; dry weight is the more accurate measure.
- The 10 per cent law: each trophic level receives just a tenth of the energy held by the level below it.
- This limits the number of levels in a grazing food chain: producer, herbivore, primary carnivore, secondary carnivore.
09 · NCERT §12.5
Ecological pyramids
- Food or energy relationships between trophic levels, drawn as bars, give a pyramid shape: broad at the base, narrow at the apex.
- The base stands for producers (first trophic level) and the apex for the top consumers.
- Three kinds are usually studied: the pyramid of number, of biomass and of energy.
- In a grassland, the pyramid of number shows only three top carnivores supported by the production of nearly 6 million plants.
- The pyramid of biomass usually falls sharply at higher trophic levels.
- In most ecosystems all three pyramids are upright: producers outnumber and outweigh herbivores, which outnumber and outweigh carnivores.
- Exceptions: counting the insects on one big tree, and then the small and larger birds that depend on them, gives an inverted pyramid of number.
- The pyramid of biomass in the sea is generally inverted: a small standing crop of phytoplankton supports a much larger biomass of zooplankton and fishes.
- The pyramid of energy is always upright and can never be inverted, because some energy is lost as heat at every transfer. Producers convert only about 1 per cent of the sunlight available to them into NPP.
10 · NCERT §12.5
Limits of trophic levels and pyramids
- Any calculation of energy, biomass or number must include all the organisms at a trophic level; a few individuals cannot support a generalisation.
- A trophic level describes a role, not a species. A single species can work at two or more trophic levels in one ecosystem at once.
- Feeding on seeds, fruits and peas, a sparrow acts as a primary consumer; feeding on insects and worms, it acts as a secondary consumer.
- Ecological pyramids do not account for a species that belongs to two or more trophic levels.
- They assume a simple food chain, which almost never exists in nature, and cannot show a food web.
- Saprophytes are left out of ecological pyramids altogether, despite their vital role in the ecosystem.
Revise
Must-know facts
- 01An ecosystem is a functional unit of nature: organisms interacting with each other and with their physical environment.
- 02Terrestrial: forest, grassland, desert. Aquatic: pond, lake, wetland, river, estuary. Man-made: crop field, aquarium.
- 03Structural features: species composition and stratification (trees on top, then shrubs, then herbs and grasses).
- 04Four functional aspects: productivity, decomposition, energy flow, nutrient cycling.
- 05Pond: phytoplankton and plants produce; zooplankton consume; fungi, bacteria and flagellates decompose.
- 06Productivity units: g m⁻² yr⁻¹ or kcal m⁻² yr⁻¹.
- 07GPP − R = NPP; NPP is what heterotrophs can use. Secondary productivity is the rate of new organic matter formed by consumers.
- 08Biosphere NPP about 170 billion tons dry weight a year; oceans, about 70 per cent of the surface, give only 55 billion tons.
- 09Decomposition steps: fragmentation, leaching, catabolism, humification, mineralisation.
- 10Lignin and chitin slow decomposition; nitrogen and sugars speed it; warm and moist favour it, cold and anaerobiosis inhibit it.
- 11The sun powers all ecosystems except deep-sea hydrothermal ones.
- 12PAR is under 50 per cent of incident sunlight; plants capture 2-10 per cent of PAR.
- 13Energy flow is unidirectional: sun → producers → consumers.
- 14Grazing food chain: grass → goat → man. Detritus food chain starts with dead organic matter.
- 15Aquatic: GFC is the main energy channel. Terrestrial: more energy flows through the DFC.
- 1610 per cent law: only 10 per cent of energy passes to the next trophic level.
- 17Standing crop: living biomass or number per unit area at a trophic level; dry weight is more accurate.
- 18Pyramid of number inverted for a single tree; pyramid of biomass inverted in the sea; pyramid of energy always upright.
- 19A sparrow is a primary consumer eating seeds and a secondary consumer eating insects.
- 20Pyramids ignore food webs, species at two levels, and saprophytes.
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Common traps
Glossary
Key terms
- Ecosystem
- A functional unit of nature where organisms interact with each other and their physical surroundings.
- Stratification
- Vertical layering of species at different levels, such as trees, shrubs and herbs in a forest.
- Species composition
- The identity and number of plant and animal species in an ecosystem.
- Primary production
- Biomass made per unit area over a period by plants in photosynthesis.
- Gross primary productivity (GPP)
- Rate of production of organic matter in photosynthesis.
- Net primary productivity (NPP)
- GPP minus respiration losses; the biomass available to heterotrophs.
- Secondary productivity
- Rate of formation of new organic matter by consumers.
- Detritus
- Dead plant and animal remains, including fecal matter; the raw material of decomposition.
- Detritivore
- An organism such as the earthworm that breaks detritus into smaller pieces.
- Humus
- Dark, amorphous, colloidal matter that resists decay and stores nutrients.
- Mineralisation
- Release of inorganic nutrients as microbes break down humus.
- PAR
- Photosynthetically active radiation, under 50 per cent of incident sunlight.
- Saprotroph
- A decomposer that feeds by breaking down dead organic matter.
- Food web
- Interconnected food chains, including links between grazing and detritus chains.
- Trophic level
- An organism's place in a food chain based on its source of food.
- Standing crop
- Mass or number of living organisms at a trophic level at a given time.
- 10 per cent law
- Only about 10 per cent of energy passes from one trophic level to the next.
- Ecological pyramid
- Bars showing number, biomass or energy at successive trophic levels.